SNP芯片开发及其在畜禽遗传育种中的应用研究进展

doi:10.19543/j.cnki.1673-4556.20260428.005

Abstract

Abstract:

Single nucleotide polymorphism (SNP) chips are high-throughput tools for detecting genetic variations in the genome. With the advantages of high detection efficiency and accurate genotyping, they exhibit broad application prospects in livestock and poultry breeding. Researchers have developed a variety of SNP chips, which provide important genome‑wide analytical methods for genetic breeding, breed improvement, and germplasm conservation of livestock and poultry.This paper reviews the development of SNP chips and their applications in key fields such as germplasm identification and genetic diversity analysis. Studies have shown that with the increase in SNP chip density from low to medium and high, their ability to capture genetic variations at the genome‑wide level is continuously improved, providing a reliable approach for accurate breed identification, genetic purity evaluation, and breed admixture detection.In addition, genome‑wide association study (GWAS) and genomic selection (GS) based on SNP chips can rapidly identify key genes and molecular markers associated with economically important traits including growth, reproduction, and disease resistance, thereby significantly improving the accuracy of selection and the efficiency of breeding.In conclusion, SNP chips have become an important platform for genomic research and molecular breeding in livestock and poultry.

Key words: SNP chip, Breeding, Livestock and poultry

CLC Number:

Ruiqi Cheng, Huaqian Zhou, Hua Yang, Yonglin Yang, Qian Yu, Wenzhe Zhang, Yan Chen, Zongsheng Zhao, lei Cui, Chunping Ma. Research progress on SNP chip development and its application in livestock and poultry genetic breeding[J].Chinese Livestock and Poultry Breeding, 2026, 22(5): 61-69.

Figures/Tables 5

Fig. 1

Table 1

Table 2

Table 3

Table 4

References 44

[1]	陈信佑, 吴平先, 巫小倩, 等. 基于SNP芯片的荣昌猪全基因组选择信号分析[J]. 中国畜牧兽医, 2025, 52(11): 5021-5029.
	CHEN X Y, WU P X, WU X Q, et al. Analysis of selection signatures for Rongchang pigs across the whole genome based on SNP chip[J]. China Animal Husbandry & Veterinary Medicine, 2025, 52(11): 5021-5029.
[2]	潘章源, 贺小云, 刘秋月, 等. 全基因组测序(WGS)在畜禽群体进化和功能基因挖掘中的应用[J]. 农业生物技术学报, 2016, 24(12): 1945-1954.
	PAN Z Y, HE X Y, LIU Q Y, et al. Application of whole genome sequencing (WGS) in population evolution and gene discovery of domestic animals[J]. Journal of Agricultural Biotechnology, 2016, 24(12): 1945-1954.
[3]	李涵, 梁胜利, 刘婷婷, 等. 基因组选择育种在畜牧育种中的应用研究进展[J]. 现代农业科技, 2025(13): 126-129.
	LI H, LIANG S L, LIU T T, et al. Research progress on the application of genome selection breeding in animal husbandry breeding[J]. Modern Agricultural Science and Technology, 2025(13): 126-129.
[4]	刘继强, 郝晓东, 武丽娜, 等. 全基因组SNP分型技术在畜禽遗传育种研究中的应用[J]. 畜牧兽医学报, 2022, 53(12): 4123-4137.
	LIU J Q, HAO X D, WU L N, et al. Application of whole genome SNP genotyping technology in livestock and poultry genetics and breeding[J]. Acta Veterinaria et Zootechnica Sinica, 2022, 53(12): 4123-4137.
[5]	马丽娜, 刘永进, 王锦, 等. 芯片技术在畜禽育种中的应用研究进展[J]. 中国畜牧兽医, 2020, 47(1): 98-106.
	MA L N, LIU Y J, WANG J, et al. Research advance on application of chip technology in livestock and poultry breeding[J]. China Animal Husbandry & Veterinary Medicine, 2020, 47(1): 98-106.
[6]	郭超凡, 陈素娟, 张珂珂, 等. SNP芯片技术在生猪育种中的应用研究进展[J]. 中国畜牧杂志, 2025, 61(11): 80-87.
	GUO C F, CHEN S J, ZHANG K K, et al. Research progress on the single nucleotide polymorphism array in pig breeding[J]. Chinese Journal of Animal Science, 2025, 61(11): 80-87.
[7]	池晓菲, 舒庆尧. 生物芯片技术的原理与应用[J]. 遗传, 2001, 23(4): 370-374.
	CHI X F, SHU Q Y. The principle and application of bio - chip technology[J]. Hereditas, 2001, 23(4): 370-374.
[8]	王有栋, 曹志平, 李玉茂, 等. SNP芯片技术原理及其在鸡遗传育种中的应用[J]. 畜牧兽医学报, 2025, 56(9): 4165-4175.
	WANG Y D, CAO Z P, LI Y M, et al. The principle of SNP chip technology and its application in chicken genetic breeding[J]. Acta Veterinaria et Zootechnica Sinica, 2025, 56(9): 4165-4175.
[9]	FU Q, LI Y C, LV Y H, et al. Fluorescent DNA-polymer enables signal amplification and precise protein localization in immunofluorescence imaging[J]. Analytical Chemistry, 2025, 97(29): 15844-15854.
[10]	PEASE A C, SOLAS D, SULLIVAN E J, et al. Light-generated oligonucleotide arrays for rapid DNA sequence analysis[J]. Proceedings of the National Academy of Sciences of the United States of America, 1994, 91(11): 5022-5026.
[11]	MCGALL G, LABADIE J, BROCK P, et al. Light-directed synthesis of high-density oligonucleotide arrays using semiconductor photoresists[J]. Proceedings of the National Academy of Sciences of the United States of America, 1996, 93(24): 13555-13560.
[12]	李宁, 丁向东, 张勤, 等. 靶向测序基因型检测 (GBTS) 技术及其应用 [J]. 中国农业科学, 2022, 55 (18): 3501-3512.
	LI N, DING X D, ZHANG Q, et al. Targeted sequencing-based genotyping (GBTS) technology and its applications[J]. Scientia Agricultura Sinica, 2022, 55(18): 3501-3512.
[13]	VLAIC B A, VLAIC A, RUSSO I R, et al. Analysis of genetic diversity in Romanian carpatina goats using SNP genotyping data[J]. Animals, 2024, 14(4): 560.
[14]	张猛, 李亮, 陈洁, 等. 液相芯片技术在动物遗传育种中的应用与发展 [J]. 中国畜牧杂志, 2025, 61 (10): 1-9.
	ZHANG M, LI L, CHEN J, et al. Application and development of liquid-phase chip technology in animal genetics and breeding[J]. Chinese Journal of Animal Science, 2025, 61(10): 1-9.
[15]	郭应威. 绵羊40K液相基因芯片的开发与应用[D]. 杨凌: 西北农林科技大学, 2022.
	GUO Y W. Development and application of a 40K liquid-phase gene chip in sheep[D]. Yangling: Northwest A&F University,2022.
[16]	贺喜, 马豪杰, 刘会超, 等. “湘芯一号”肉鸡60K液相育种芯片的设计及基因组预测效果[J]. 湖南农业大学学报(自然科学版), 2024, 50(6): 1-9.
	HE X, MA H J, LIU H C, et al. Design and genomic prediction effects of the“Xiangxin No. 1”broiler 60K liquid phase breeding chip[J]. Journal of Hunan Agricultural University (Natural Sciences), 2024, 50(6): 1-9.
[17]	颉满斌. 兰州大学研发“华羊芯”绵羊育种芯片[N]. 科技日报, 2024-01-02(6).
	XIE M B. Lanzhou University develops the “Huayangxin” sheep breeding chip[N]. Science and Technology Daily, 2024-01-02(6).
[18]	中国农网.白泽一号, 中国绵羊育种之芯, 赋能畜禽种业振兴[EB/OL]. , 2023-12-14.
	China Agricultural News Network. Baize-1, the core of Chinese sheep breeding, empowering the revitalization of livestock and poultry seed industry[EB/OL]. , 2023-12-14.
[19]	李隐侠, 牙生江·那斯尔, 赛里克·都曼, 等. SNP芯片评估柯尔克孜羊群体遗传多样性和遗传结构[J]. 畜牧兽医学报, 2023, 54(2): 572-583.
	LI Y X, YA S J S, K·D SAI L, et al. Evaluation of genetic diversity and genetic structure in kirgiz sheep population based on SNPs chip[J]. Acta Veterinaria et Zootechnica Sinica, 2023, 54(2): 572-583.
[20]	杨志勇, 孙一斐, 白凤庭, 等. 阿旺绵羊群体结构及毛色性状关联分析[J]. 家畜生态学报, 2024, 45(11): 7-13.
	YANG Z Y, SUN Y F, BAI F T, et al. Association between population structure and coat color traits in awang sheep[J]. Acta Ecologae Animalis Domastici, 2024, 45(11): 7-13.
[21]	刘佳森, 张莉, 李蕴华, 等. 苏尼特羊拷贝数变异的基因组分布特征研究[J]. 中国畜牧兽医, 2013, 40(10): 173-178.
	LIU J S, ZHANG L, LI Y H, et al. Genomic characteristics of copy number variations in sunite sheep[J]. China Animal Husbandry & Veterinary Medicine, 2013, 40(10): 173-178.
[22]	LI X, YANG J, SHEN M, et al. Whole-genome resequencing of wild and domestic sheep identifies genes associated with morphological and agronomic traits[J]. Nature Communications, 2020, 11: 2815.
[23]	蒋立斌, 张云生, 杨华, 等. 绵羊FecB基因非探针法HRM分型技术条件优化研究[J]. 畜牧与兽医, 2014, 46(5): 55-58.
	JIANG L B, ZHANG Y S, YANG H, et al. Optimization of technical conditions for non-probe HRM typing of FecB gene in sheep[J]. Animal Husbandry & Veterinary Medicine, 2014, 46(5): 55-58.
[24]	MEUWISSEN T H, HAYES B J, GODDARD M E. Prediction of total genetic value using genome-wide dense marker maps[J]. Genetics, 2001, 157(4): 1819-1829.
[25]	KIJAS J W, LENSTRA J A, HAYES B, et al.Genome-Wide Analysis of the World's Sheep Breeds Reveals High Levels of Historic Mixture and Strong Recent Selection[J]. PLoS Biology, 2012, 10(2): e1001258.
[26]	BENEDETTI DEL RIO E, MANCIN E, ORSI M, et al. Genomic characterisation of local sheep breeds of the Eastern Alps[J]. Italian Journal of Animal Science, 2025, 24(1): 2515271.
[27]	刘坤. 西农大团队研发出奶绵羊育种专用芯片[N]. 陕西科技报, 2023-07-11(1).
	LIU K. Team from Northwest A&F University develops special chip for dairy sheep breeding[N]. Shaanxi Science and Technology News, 2023-07-11(1).
[28]	田可川, 付雪峰. 一种基于靶向捕获测序的超细型细毛羊 10K 液相芯片及其应用[EB/OL]. , 2024-6-3.
	TIAN K C, FU X F.A 10 K liquid chip for ultra-fine wool sheep based on target capture sequencing and its application[EB/OL]. , 2024-6-3.
[29]	安徽农业大学动物科技学院. 羊种质资源创新与利用团队成功创制我省首款羊育种专制基因芯片[EB/OL]. , 2024-11-22.
	College of Animal Science and Technology, Anhui Agricultural University. The sheep germplasm resource innovation and utilization team successfully creates Anhui's first special gene chip for sheep breeding[EB/OL]. , 2024-11-22.
[30]	兰蓉, 朱兰, 杨红远, 等. 基于SNP芯片的云上黑山羊遗传结构分析[J]. 中国畜牧兽医, 2019, 46(2): 480-488.
	LAN R, ZHU L, YANG H Y, et al. Analysis of genetic structure of Yunshang black goat based on SNP chip[J]. China Animal Husbandry & Veterinary Medicine, 2019, 46(2): 480-488.
[31]	河南牧业经济学院. 我校牵头研发的 “豫羊牧芯” 河南省首个肉羊育种芯片重磅发布[EB/OL]. , 2025-1-11.
	Henan University of Animal Husbandry and Economy. The university takes the lead in developing "Yuyang Muxin", the first meat sheep breeding chip in Henan Province officially released[EB/OL]. , 2025-1-11.
[32]	BAGNATO A, SCHIAVINI F, STRILLACCI M G, et al. Identification and validation of copy number variants in Italian Brown Swiss dairy cattle using Illumina Bovine SNP50 Beadchip[J]. Italian Journal of Animal Science, 2015, 14: 3900.
[33]	CREPALDI M, BIFFANI S, DALL'OLIO S, et al. Genomic approach to manage genetic variability in dairy farms[J]. Italian Journal of Animal Science, 2023, 32(1): 117-126.
[34]	王元清, 王泽昭, 朱波, 等. 不同芯片密度对华西牛重要经济性状基因组评估准确性的影响[J]. 畜牧兽医学报, 2025, 56(2): 591-602.
	WANG Y Q, WANG Z Z, ZHU B, et al. Comparison of prediction accuracy of genomic selection for economically important traits in Huaxi cattle based on different chip densities[J]. Acta Veterinaria et Zootechnica Sinica, 2025, 56(2): 591-602.
[35]	吴江鸿. 蒙古牛 10K 液相育种芯片 “相牛 1 号” 的研发与应用 [J]. 内蒙古民族大学学报 (自然科学版), 2025, 46 (2):F0003 - F0003.
	WU J H. Development and application of the Mongolian cattle 10K liquid-phase breeding chip “XiangNiu No.1”[J]. Journal of Inner Mongolia Minzu University (Natural Science Edition), 2025, 46(2): F0003-F0003.
[36]	JIA X, CHEN S, ZHOU H, et al. Copy number variations identified in the chicken using a 60K SNP BeadChip[J]. Animal Genetics, 2013, 44(3): 276-284.
[37]	WOLC A, ARANGO J, O'SULLIVAN N P, et al. Genome-wide association study for feed efficiency in broilers [J]. Journal of Animal Science, 2013, 91 (10):4753-4759.
[38]	王宇哲, 刘冉冉, 赵桂苹, 等.鸡 55K SNP 芯片的设计与应用评估 [J]. 中国家禽, 2018, 40(15):1-6.
	WANG Y Z, LIU R R, ZHAO G P, et al. Design and application evaluation of a 55K SNP chip for chickens[J]. China Poultry, 2018, 40(15): 1-6.
[39]	国家家养动物种质资源库. 肉鸡基因组育种技术迭代升级 —“京芯一号 mini”育种芯片发布[EB/OL]. , 2024-09-25.
	National Animal Germplasm Resource Bank. Iterative upgrade of broiler genomic breeding technology - launch of "Jingxin No.1 Mini" breeding chip[EB/OL]. , 2024-09-25.
[40]	ZHANG Z P, XING S Y, QIU A, et al. The development of a porcine 50K SNP panel using genotyping by target sequencing and its application[J]. Journal of Integrative Agriculture, 2025, 24(5): 1930-1943.
[41]	中国科学院亚热带农业生态研究所. 湘猪育种第一“芯”:64K cGPS 液相芯片研发成功[EB/OL]. , 2023-04-14.
	Institute of Subtropical Agriculture, Chinese Academy of Sciences. The first "core" for Hunan pig breeding: successful development of 64K cGPS liquid-phase chip[EB/OL]. , 2023-04-14.
[42]	中国农科新闻网. 安徽农大联合博睿迪首款种猪育种专用基因芯片研发成功[EB/OL]. , 2023-02-22.
	China Agricultural Science News Network. Anhui Agricultural University and Boruidi successfully developed the first sp
	ecial gene chip for pig breeding[EB/OL]. , 2023-02-22.
[43]	刘佳惠子, 李芳. 核心技术指标世界领先 “中芯一号”猪育种芯片实现全面国产化[J]. 猪业观察, 2024(5): 7.
	LIU J, LI F. The core technical indicators of the world-leading “SMIC-1” pig breeding chip are fully localized[J]. Swine Industry Outlook, 2024(5): 7.
[44]	拉索生物. 拉索生物、温氏股份共同推出猪育种固相基因芯片[EB/OL]. , 2024-10-16.
	Bio Lasso. Lasso Bio and Wens Foodstuff Co., Ltd. jointly launch solid-phase gene chip for pig breeding[EB/OL]. , 2024-10-16.

Metrics

Viewed

Full text

Abstract

Cited

Shared

Discussed

Comments

Recommended 0

No Suggested Reading articles found!

年份Year	芯片名称 Chip name	密度Density/K	类型 Types	研发单位或团队 Research institution or team	用途 Purpose	参考文献References
2016	BovineSNP50 v3 DNA Analysis BeadChip	50	固相芯片	Illumina公司与美国农业部农业研究服务局、密苏里大学和阿尔伯塔大学	全基因组选择、数量性状位点鉴定、个体遗传价值评估等	[32]
2016	GGP Bovine 100K	100	固相芯片	Neogen公司	奶牛和肉牛的遗传评估、选育等	[33]
2023	Cattle110K	110	液相芯片	中国农业科学院北京畜牧兽医研究所李俊雅团队	肉牛的遗传改良	[34]
2025	相牛1号	10	液相芯片	内蒙古民族大学吴江鸿教授团队联合内蒙古牧业技术推广中心	基因检测和遗传资源鉴定等	[35]

年份Year	芯片名称 Chip name	密度Density/K	类型Types	研发单位或团队 Research institution or team	用途 Purpose	参考文献References
2011	Chicken 60K SNP Beadchip	60	固相芯片	Illumina公司	肉鸡相关性状的遗传分析	[36]
2011	Illumina Chicken 60K SNP BeadChip	60	固相芯片	瓦赫宁根大学、华盛顿大学医学院等机构	鸡的全基因组关联研究和基因组选择育种等。	[37]
2017	京芯一号	50	液相芯片	中国农业科学院北京畜牧兽医研究所赵桂苹团队	肉鸡全基因组选择	[38]
2024	京芯一号mini	10	液相芯片	中国农业科学院北京畜牧兽医研究所赵桂苹团队	肉鸡的基因组育种	[39]

年份Year	芯片名称 Chip name	密度Density/K	类型 Type	研发单位或团队 Research institution or team	用途 Purpose	参考文献References
2021	猪50 K液相芯片	50	液相芯片	中国农业大学丁向东研究团队	遗传分析和分子育种	[40]
2022	湖南地方猪 60 K SNP芯片	60	液相芯片	中国科学院亚热带农业生态研究所印遇龙院士团队与华智生物公司	对湖南地方猪进行基因型精准鉴定评估及重要性状基因的挖掘利用	[41]
2023	皖芯1号	50	液相芯片	安徽农业大学猪遗传育种创新团队与博瑞迪生物技术有限公司	地方猪的基因精准鉴定及重要性状基因的挖掘利用等	[42]
2023	皖芯2号	50	液相芯片	安徽农业大学猪遗传育种创新团队与博瑞迪生物技术有限公司	瘦肉型种猪全基因组关联分析和基因组选择育种	[42]
2024	中芯一号	100	固相芯片	温氏食品集团股份有限公司与苏州拉索生物芯片科技有限公司	家猪基因组育种	[43]
2024	PorcineWENS 100K	100	固相芯片	温氏食品集团股份有限公司与苏州拉索生物芯片科技有限公司	进行基因组选择，提高育种效率	[44]

Research progress on SNP chip development and its application in livestock and poultry genetic breeding

RichHTML

PDF (PC)

Abstract

Cite this article

share this article

Figures/Tables 5

References 44

Related Articles 14

Metrics

Comments

Recommended 0

年份Year	芯片名称 Chip name	密度Density/K	类型Type	研发单位或团队 Research institution or team	用途 Purpose	参考文献References
2009	Illumina Ovine SNP50 K BeadChip	50	固相芯片	Illumina公司与国际绵羊基因组联盟（ISGS）	绵羊基因分型、群体遗传结构分析、筛选性状相关分子标记和基因等	[25]
2020	GeneSeek Genomic Profiling (GGP) Ovine 50 K	50	固相芯片	Neogen公司	羊的亲子鉴定、筛选不良突变	[26]
2023	华羊芯	45	液相芯片	兰州大学王维民教授团队	绵羊全基因组选择育种、经济性状基因定位、遗传多样性评估等	[17]
2023	奶绵羊20 K育种芯片	20	液相芯片	西北农林科技大学宋宇轩教授团队	奶绵羊全基因组选择育种、建立种羊基因身份证、分子标记辅助选择等	[27]
2023	白泽一号	40	液相芯片	中国农业大学韩红兵教授团队	绵羊遗传多样性评估、遗传图谱构建及基因定位、种质资源鉴定、亲缘鉴定、全基因组关联分析、全基因组选择等	[18]
2024	皖羊1号	50	液相芯片	安徽农业大学凌英会教授联合北京康普森农业科技有限公司	山羊全基因组选择育种、全基因组关联分析、血统渗入及亲缘关系鉴定等	[29]
2024	“云上黑山羊”50 K育种芯片	50	液相芯片	云南省畜牧兽医科学院洪琼花研究员团队	云上黑山羊全基因组选择育种、亲缘关系鉴定等	[30]
2024	超细型细毛羊10 K液相基因芯片	10	液相芯片	山东省农业科学院田可川研究员团队和新疆畜牧科学院付雪峰研究员团队	羊基因分型、分子标记辅助育种、遗传多样性分析、种羊基因身份证研究	[28]
2025	豫羊牧芯	10	固相芯片	河南牧业经济学院权凯教授团队	预测羊的体重、生长速度、饲料转化率	[31]

[1]	Duanyang Ren, Wenxia Li, Xinpei Wang, Xi Wang. Population genetic structure analysis and molecular mating of Jinnan cattle based on genome-wide SNP loci [J]. Chinese Livestock and Poultry Breeding, 2026, 22(6): 72-80.
[2]	Hai Wang, Daoning Yu, Chun Zhang, Qianqian Chu, Junjun Zhao, Daiyang Zhang, Congyong Wang. Research on innovation and industrialization of breeding technology of Huaxi cattle [J]. Chinese Livestock and Poultry Breeding, 2026, 22(6): 38-44.
[3]	Lisheng Miao, Congyong Wang, Yang Cao, Yumin Zhao. Development of the beef cattle seed industry and genetic improvement pathways [J]. Chinese Livestock and Poultry Breeding, 2026, 22(6): 25-30.
[4]	Fengjuan Liu, Tiecheng Wu, Junyang Liu, Tao Wang, Leqi Yi, Yulin Gao, Xingang Yan, Bin Liu. Research progress on breeding and selection techniques for domestic animal genomes [J]. Chinese Livestock and Poultry Breeding, 2026, 22(5): 70-78.
[5]	Cuiping Cao. Analysis of factors contributing to Denmark's high PSY in swine production and implications [J]. Chinese Livestock and Poultry Breeding, 2026, 22(5): 7-10.
[6]	Tiantian Wang, Jiahao Shao, Wenmiao Duan, Jianing Lu, Jianhua Zeng, Xiaohong Liu, Xiaolong Yuan. Research progress on genetic diversity of coat color in livestock genetic breeding [J]. Chinese Livestock and Poultry Breeding, 2026, 22(5): 108-115.
[7]	Bai Hongjie. Information Management and Brand Building of Seed Enterprises in the Background of Strong Agricultural Province—Taking Henan NongKe Pig Breeding Technology Co., Ltd. as an example [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(8): 106-110.
[8]	Zhao Xiaoqiang, Zhang Yuanqing. Thoughts on Developing Rural Beef Cattle Production: A Case Study of A Village, Wenshui County, Shanxi Province [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(7): 146-150.
[9]	Wang Ting, Zhu Bo, Wang Zezhao, Chen Yan, Xu Lingyang, Zhang Lupei, Gao Xue, Gao Huijiang, CaoYang, Zhao Yumin, Li Junya. History of the Development of Domestic and Foreign Breed Bull Auctions [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(7): 128-134.
[10]	Wang Honghao, Wang Xi, Che Leijie, Lv Huaixin, Yan Qi, Ming Shiqing. Conservation, Breeding and Utilization of Jinnan Cattle [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(7): 97-101.
[11]	Wang Honghao, Jin Guang, Li Bo, Cheng Jing, Zhang Dandan, Liu Yanjie, Niu Xiaoyan, Li Xi, Zhang Yuanqing. Application of Quantitative Genetics in Beef Cattle Breeding [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(7): 74-79.
[12]	Dou Yuting, Zhang Yuanxu, Li Jing, Wang Zezhao, Chen Yan, Xu Lingyang, Zhang Lupei, Gao Huijiang, Gao Xue, Li Junya, Li Hui, Zhu Bo. Research Progress on Hornless Traits of Beef Cattle [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(7): 61-67.
[13]	Wu Yu, Zhou Di, Chen Kun, Jiang Gui rong, Wang Yan, Yang Rong, AO Ye, Fang Hua, Wang She. Research Progress on Candidate Genes for Semen Quality in Boars [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(6): 105-112.
[14]	Zhang Geyang, Lv Shijie, Zhu Xiaoting, Zhai Yaying, Zhang Zhihao, Shi Qiaoting, Zhang Zijing, Qu Junfeng, Chen Fuying, Wang Eryao. Application of Liquid Chip Technology in Animal Genetics and Breeding [J]. The Chinese Livestock and Poultry Breeding, 2023, 19(6): 24-28.